Reptiles - Part V - The Zoology of the Voyage of H.M.S Beagle: Under the Command of Captain Fitzroy - During the Years 1832 to 1836
By Charles Darwin and Thomas Bell
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Charles Darwin
Charles Darwin (1809–19 April 1882) is considered the most important English naturalist of all time. He established the theories of natural selection and evolution. His theory of evolution was published as On the Origin of Species in 1859, and by the 1870s is was widely accepted as fact.
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Reptiles - Part V - The Zoology of the Voyage of H.M.S Beagle - Charles Darwin
REPTILES.
TRIBUS—EUNOTES. Bibron.
FAMILIA—TROPIDURIDÆ. Mihi.
TROPIDOLEPIDIENS. Bibr.
GENUS—Tropidurus. (In part.) Weigmann.
PROCTOTRETUS. Bibr.
CHARACTER GENERICUS.—Pori femorales nulli. Pori prœ-anales in maribus tantum. Crista dorsalis nulla. Dentes palatini. Squamœ imbricatæ; dorsales carinatæ.
THE genus which I take the present favourable opportunity to illustrate, formed a section or sub-genus of the genus Tropidurus, according to Weigmann, who, however, was acquainted with two species only; Pr. Chilensis and Pr. nigromaculatus. Of the varieties of the former of these, that author has made no less than three species; but these have been very properly reduced by M. Bibron to one only. The last-named excellent erpetologist described several additional species, which had been brought from Chile, by M. D’Orbigny, and others; and I received, some years since, from Capt. King, three or four species which were found by him in the same locality, in the course of his well-known survey. The number of species altogether, hitherto known, amounts to ten; to which I have now the opportunity of adding four entirely new, forming part of the interesting collection of Reptilia made by Mr. Darwin. One of them, Pr. Kingii, was already in my collection, amongst those which were given me by Capt. King. The genus, therefore, of which, but lately, two species only were known, now consists of fourteen; and it is highly probable that more may yet be obtained by more prolonged and extensive investigation in the same districts.
Thus of the species now described two were known to Weigmann, and described by him, namely, Chilensis, and nigromaculatus. I received from Capt. King, Chilensis, pictus, cyanogaster, Kingii, Fitzingerii, and, from other sources, Chilensis, pictus, Weigmannii. Bibron describes the whole of these, excepting Kingii, and in addition to them tenuis, pectinatus, signifer. In Mr. Darwin’s collection are found all those described by Bibron, excepting signifer; and in addition Kingii now first described, and the following species never before observed, namely, Darwinii, gracilis, Bibronii.
Although the form of the whole of the species much more nearly approximates that of the Agamidæ than most others, they are far removed from that family by several important characters, which it is unnecessary here to detail.
PROCTOTRETUS CHILENSIS.
PLATE I.—FIG. 1.
Auribus margine anteriore dentato; collo non plicato; squamis dorsi magnis, rhombeis; acutè carinatis; serie unicâ squamarum supralabialium.
Calotes Chilensis, Less. et Garn. Voy. de la Coquille, Zool. Rep. t. i. f. 2.
Tropidurus Chilensis, Weigm. Act. Acad. Cæs. Leop. Carol. Nat. cur. xvii. pp. 233. 268.
Proctotretus Chilensis, Bibr. Hist. Nat. des Rep. IV. p. 269.
Sun. Tropidurus nitidus, Weigm. 1. c. p. 234, t. xvii. f. 2.
Var. Tr. olivaceus, Ib. 1. c. p. 268.
Habitat, Guasco in Northern Chile.
DESCRIPTION.—Head short and broad, rostrum rounded, obtuse. Scales of the head large, and slightly raised, separated by distinct grooves. Superciliary ridge strongly marked, forming a distinct carina, composed of five or six narrow, elongated, obliquely imbricated scales. Nostrils large, nearly round. A single scries of narrow scales between those of the upper lip and the orbit. Scales of the temples imbricated, rhomboidal and carinated. The opening of the ear oval, rather large, furnished anteriorly with three or four projecting scales, of which the upper one is the largest. The neck is short, robust and round, and without any lateral fold; in which it differs from every other species of the genus. The trunk is thick, rounded on the back and sides, flattened beneath, diminishing toward either extremity. The tail is ordinarily almost twice as long as the body, thick at its origin, and tapering regularly to the extremity, nearly round, excepting near the base, where it is slightly quadrilateral. The anterior feet when placed against the sides, extend backwards little more than mid-way between the shoulder and the groin; the posterior ones, stretched forward, reach the arm-pit.
The scales of the whole upper and lateral parts of the body, tail, and limbs, are loosely imbricated, large, rhomboidal, and furnished with an elevated carina, terminating in an acute point: those of the under parts are large, smooth, and obtuse.
This is one of the most beautiful species of the genus. The general form is robust and solid, forming a remarkable contrast with most other species of the genus. The surfaee is beautifully relieved by the fine, large and prominent scales, which are ranged in rows of perfect regularity, of which there are about eighteen on the back and