Cacti of California
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This title is part of UC Press's Voices Revived program, which commemorates University of California Press’s mission to seek out and cultivate the brightest minds and give them voice, reach, and impact. Drawing on a backlist dating to 1893, Voices Revived
E. Yale Dawson
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Cacti of California - E. Yale Dawson
California Natural History Guides: 18
THE CACTI
OF
CALIFORNIA
BY
E. YALE DAWSON
WITH PHOTOGRAPHS BY DON SKINNER
UNIVERSITY OF CALIFORNIA PRESS
BERKELEY, LOS ANGELES, LONDON
CALIFORNIA NATURAL HISTORY GUIDES
ARTHUR C. SMITH, GENERAL EDITOR
UNIVERSITY OF CALIFORNIA PRESS
BERKELEY AND LOS ANGELES, CALIFORNIA
UNIVERSITY OF CALIFORNIA PRESS, LTD.
LONDON, ENGLAND
© 1966 BY THE REGENTS OF THE UNIVERSITY OF CALIFORNIA
LIBRARY OF CONGRESS CATALOG CARD NUMBER: 65-25843
PRINTED IN THE UNITED STATES OF AMERICA
FOURTH PRINTING, 1982
ISBN: 0-520-02990-0 (CLOTHBOUND)
0-520-00299-7 (PAPERBOUND)
CONTENTS
CONTENTS
INTRODUCTION
WHAT ARE CACTI?
HOW TO STUDY CALIFORNIA CACTI
WHERE TO FIND THEM
KEY TO THE GENERA AND SUBGENERA OF CALIFORNIA CACTI
THE CACTI OF CALIFORNIA
OPUNTIA
CYLINDROPUNTIA: THE CHOLLAS
PLATYOPUNTIA: THE PRICKLY PEARS
Giant Cactus; Saguaro (Carnegiea gigantea)
Velvet Cactus (Bergerocactus emoryi)
Hedgehog Cactus; Strawberry Cactus (Echinocereus engelmannii)
Desert Barrel Cactus (Ferocactus acanthodes)
Nigger Heads (Echinocactus polycephalus)
Pigmy Barrel Cactus (Echinomastus johnsonii)
Long-spined Fishhook Cactus (Sclerocactus polyancistrus)
Desert Pincushion Cactus (Coryphantha deserti)
Coastal Fishhook Cactus; Nipple Cactus (Mammillaria dioica)
Cork-seeded Fishhook Cactus (Phellosperma tetrancistra)
CACTUS BOOKS AND CLUBS
GLOSSARY
INDEX
INTRODUCTION
The study of cacti in California has had a long history. Except for the prickly pear of the Atlantic states, known to Linnaeus, and four species from the upper Missouri River described in 1814 as a result of the exploration of the Louisiana Purchase, the first cacti in this country to be known to science came from San Diego. In 1834 Thomas Nuttall, pioneer of western American botany, discovered in the village of San Diego, then little more than the Mexican mission and its Indians, "Echinocactus viridescens" and "C er eus californicus.,,
By 1856, through the work of George Engelmann of St. Louis on collections obtained by the Pacific Railroad Survey and the explorations of the Mexican Boundary Commission, knowledge of California cacti was spreading rapidly. By 1919, when the first volume of the monumental monograph on Cactaceae by Britton and Rose was being published, a large body of information on California cacti had accumulated, and our deserts had been extensively explored.
A number of kinds had eluded the botanists, however, and these have continued to be discovered and described down to the present day. A new publication lies before me, just received in the mail, in which a new species of California prickly pear is named. Its characters have been confused or overlooked all these years and only now are clearly recognized. Similarly, the careful exploration of our desert ranges along the Arizona and Mexican boundaries has revealed very recently the occurrence in California of species hitherto known only from these other areas. Two of these are reported for the first time in this handbook. Perhaps you may find still others.
The more we explore and the more we learn of the cacti, the less certain we become of just how many kinds there are. Most of the species that were discovered many years ago appeared to be distinct from one another, but as collections expanded and observations became more critical, we noted numerous intergradations and intermediate conditions. Thus, where three species had been named earlier, we may now recognize only one, perhaps in three major variable groups that merge. Furthermore, we are increasingly aware that evolution in the cacti is going on before our eyes, that distinct species (and even genera) in nature are occasionally producing crosses that survive and become, in fact, new species. We have sometimes hastened this process by artificial introduction of species from other areas, particularly the Mission Cactus. The resulting hybridization with native species has added many variables to original natural populations.
Some of the difficulties in recognizing species among these variable cacti are related to their widespread distribution and variation in far-flung geographic areas. As a broad
species may have a varied population in Texas, another in Arizona, and still another in California, so the California plants may scarcely resemble the Texas plants, yet may be linked by intermediates. Interpretations by different botanists of these variations have led to the use of different scientific names in designating the plants; so do not be dismayed if the name used here does not take the form that ap- ears in another book. We have chosen names that, or a local flora, seem most convenient and useful in designating the California plants. A flora of North America would take a different point of view.